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Robert Mills does not work for, consult, own shares in or receive funding from any company or organisation that would benefit from this article, and has disclosed no relevant affiliations beyond their academic appointment. The prevailing view is that animals mainly have sex to reproduce. Until recently, therefore, scientists assumed that animals were relentlessly heterosexual. This is the message conveyed by countless zoos, wildlife documentaries, books and films. But arguing against these viewpoints, biologists such as Bruce Bagemihl and Joan Roughgarden have begun putting forward evidence that animal sexuality comprises an array of behaviours, gender expressions and body types. In fact, reproduction is marginal to many species.


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The Laysan albatross is a downy seabird with a seven-foot wingspan and a notched, pale yellow beak. Every November, a small colony of albatrosses assembles at a place called Kaena Point, overlooking the Pacific at the foot of a volcanic range, on the northwestern tip of Oahu, Hawaii. Each bird has spent the past six months in solitude, ranging over open water as far north as Alaska, and has come back to the breeding ground to reunite with its mate.

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